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Olivia Neilly, who is earning a double major in English and molecular, cellular and developmental biology with a perfect 4.0 GPA, is named the college’s outstanding graduate for fall 2025

When Olivia Neilly stepped onto the University of Colorado Boulder campus four years ago, she thought she had her future mapped out.

“I really wanted to go to medical school,” she recalls. “I thought I’d keep my head in the books for four years and then move on.”

However, in pursuit of courses that would prepare her for the medical field, Neilly joined Professor��Zoe Donaldson’s neuroscience lab in the��Department of Molecular, Cellular and Developmental Biology (MCDB)—and that one experience changed everything for her.

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Olivia Neilly is the Fall 2025 College of Arts and Sciences outstanding graduate. (Photo: Julie Chiron)

“I discovered that research is not just about data—it’s about asking questions, embracing creativity and finding joy in discovery,” Neilly says. “It literally changed the trajectory of what I want to do with my life, and now I can’t imagine wanting to do anything else.”

Donaldson and��Jenny Knight, professor of molecular, cellular and developmental biology, ­became important mentors for Neilly, whom she credits with fostering creativity and curiosity in the lab. Additionally, PhD graduate Mostafa El-Kalliny helped shape her thinking about research as well as issues outside of science.

“From day one in the lab I worked with Mostafa, who shaped how to think about science—and other subjects,” she says. “Our conversations weren’t just about experiments—they were about philosophy, literature and life.”

Embracing neuroscience with a passion

For her honor’s thesis, Neilly wrote a 71-page research paper investigating how a small part of the brain called the nucleus accumbens helps animals form close social bonds, research that has potential implications for humans. Her paper specifically explored the neuroscience of social bonding on prairie voles, a small species of furry rodents.

“We study prairie voles because they form lifelong pair bonds, like humans,” Neilly explains. “My project focused on nucleus accumbens, a brain region tied to reward. I used chemogenetics (a technique that makes use of engineered proteins) to turn off specific interneurons during bond formation. When those cells were silenced, voles couldn’t form pair bonds. This suggests one cell type can influence complex social behavior, which has implications for psychiatric disorders.”

Neilly began her lab work with the voles before the start of her sophomore year and spent two summers working full time in the lab. This past summer, she completed the experiment underlying her thesis and spent the school year analyzing the data and writing. While the work was very time consuming, Neilly adds, “It never felt like a burden—I loved the process.”

In addition to that work, Neilly authored a manuscript for the scientific journal Nature Communications as well as a second manuscript currently being considered for publication.

Earning high praise from faculty

Neilly was nominated for the outstanding graduate award by Christy Fillman, chair of the MCDB Honors Committee, and Donaldson, who praised her undergraduate student for her curiosity, intellect and strong work ethic.

“I would often find Olivia in the lab at all hours, eager to contribute and learn new skills. By this time last year, she was already operating at the level of a graduate student despite being only a junior. She accomplished this while also maintaining a 4.0 GPA in two majors and maintaining her involvement in other activities, including the American Lung Cancer Society Screening Initiative,” Donaldson wrote in her letter recommending Neilly for the outstanding graduate honor. Donaldson added, “She is the most impressive undergraduate I have had the chance to mentor or interact with across institutions I have worked at.”

Neilly says receiving the outstanding graduate award is both exciting and humbling.

“My mentor (El-Kalliny) hinted that I might get nominated, but honestly, I was so focused on graduating and finishing classes that I didn’t think much about it. When I got the email and Donaldson announced the award in our lab group chat, I was really touched,” Neilly says. “I’m emotional by nature, so it meant a lot that people I respect recognized my efforts. I usually just put my head down and work, not for recognition, so this felt validating. I was proud—and excited to tell my mom first.”

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Olivia Neilly (second from left), Fall 2025 College of Arts and Sciences outstanding graduate, chats with, left to right, Daryl Maeda, interim dean of the College of Arts and Sciences; Irene Blair, dean of natural sciences; and Jennifer Fitzgerald, interim associate dean for student success. (Photo: Julie Chiron)

Balancing science and the arts

Neilly’s academic path has proven to be as unique as her research. Initially focused on MCDB, she opted to add an English degree to feed her artistic side.

“At first, I thought there was no overlap,” she recalls. “I started with MCDB for medical school or research but then added English because I missed my artistic side. Over time, I realized they overlap in surprising ways. In science, clear communication is essential—especially now, in a media environment riddled with so much misinformation."

As a creative writer and fan of modern fiction, Neilly applauds how people are pushing the boundaries of language in the same way that scientists are pushing the boundaries of scientific knowledge—celebrating the unbounded exploration of both art and science.

“Writing skills from English help me convey research effectively. Creativity is key in both fields. The best scientists are often the most creative.”

Life beyond the lab

Despite her demanding academic schedule, Neilly says she made time for extracurricular activities. She wrote articles for the online magazine Her Campus, attended film festivals and organized a lung cancer awareness event at Boulder’s historic Chautauqua Park.

She says she feels fortunate to have partaken in many cultural events offered by ���Ҵ�ý�ƽ������ and by the local community, and she encourages her fellow students to do the same, adding, “Connecting with your community matters as much as academics.”

As for any advice for incoming CU students, Neilly says, “Wherever you are, you can make the most of it if you put in the time and energy. Be willing to try new things and embrace discomfort—it’s how you grow.”

Right where she was supposed to be

Neilly says she’s grateful for her time at ���Ҵ�ý�ƽ������ and is now looking forward to what comes next as she prepares to embark on a scientific career. After graduating later this month, Neilly will join Stanford University as a research technician in Boris Heifets’ lab, where scientists study how psychoactive compounds can help treat severe psychiatric disorders.

“I’m passionate about improving mental health and social functioning, so this feels like the right next step before starting grad school,” she says.

Perhaps ironically in retrospect, Neilly says she wasn’t initially committed to attending ���Ҵ�ý�ƽ������. She earned good grades in high school and had a number of options when it came time to select a university.

“I have a long family history with CU; my mom, sister and grandfather all have ties here. At first, I thought I wanted to break the pattern, but my mom reminded me that education is what you make of it,” says Neilly, who spent much of her childhood in Aurora. “CU ended up being the best decision. I found incredible mentors and research opportunities I wouldn’t have had elsewhere.

“I don’t regret a thing. I’ve used CU to the absolute ends of what it could offer.”��

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